##### Title:

Redundancy Analysis (RDA) scores and environmental variables in the Danube - Black Sea, Elbe - North Sea, Guadalquivir - Atlantic Ocean and Po - Adriatic Sea river-to-sea continuum.



##### Description:

This dataset contains the results of Redundancy Analyses (RDAs) performed to explore relationships between environmental variables and community composition in the Danube - Black Sea, Elbe - North Sea, Gualdalquivir - Atlantic Ocean and Po - Adriatic Sea river-to-sea continuum for macrozoobenthos, fish and phytoplankton. It includes:

Environmental variable scores (loadings) on RDA axes
Sample scores (site coordinates) on the same axes

These data can be used to reconstruct ordination plots and interpret ecological gradients.

This repository contains the derived results of the MARCO-BOLO Deliverable 3.3.
The materials provided here do not include the original/raw dataset, but only processed outputs generated from it.



##### Contents:

MBO_Report_Danube_Macrozoobenthos_RDA_env_vectors.csv - RDA environmental vectors of macrozoobenthos in Danube - Black Sea river-to-sea system.
MBO_Report_Danube_Macrozoobenthos_RDA_scores.csv - RDA sample scores of macrozoobenthos in Danube - Black Sea river-to-sea system.
MBO_Report_Elbe_Fish_RDA_env_vectors.csv - RDA environmental vectors of fish in Elbe - North Sea river-to-sea system.
MBO_Report_Elbe_Fish_RDA_scores.csv - RDA sample scores of fish in Elbe - North Sea river-to-sea system.
MBO_Report_Elbe_Macrozoobenthos_RDA_env_vectors.csv - RDA environmental vectors of macrozoobenthos in Elbe - North Sea river-to-sea system.
MBO_Report_Elbe_Macrozoobenthos_RDA_scores.csv - RDA sample scores of macrozoobenthos in Elbe - North Sea river-to-sea system.
MBO_Report_Elbe_Phytoplankton_RDA_env_vectors.csv - RDA environmental vectors of phytoplankton in Elbe - North Sea river-to-sea system.
MBO_Report_Elbe_Phytoplankton_RDA_scores.csv - RDA sample scores of phytoplankton in Elbe - North Sea river-to-sea system.
MBO_Report_Guadalquivir_Fish_RDA_env_vectors.csv - RDA environmental vectors of fish in Guadalquivir - Atlantic Ocean river-to-sea system.
MBO_Report_Guadalquivir_Fish_RDA_scores.csv - RDA sample scores of fish in Guadalquivir - Atlantic Ocean river-to-sea system.
MBO_Report_Guadalquivir_Macrozoobenthos_RDA_env_vectors.csv - RDA environmental vectors of macrozoobenthos in Guadalquivir - Atlantic Ocean river-to-sea system.
MBO_Report_Guadalquivir_Macrozoobenthos_RDA_scores.csv - RDA sample scores of macrozoobenthos in Guadalquivir - Atlantic Ocean river-to-sea system.
MBO_Report_Po_Macrozoobenthos_RDA_env_vectors.csv - RDA environmental vectors of macrozoobenthos in Po - Adriatic Sea river-to-sea system.
MBO_Report_Po_Macrozoobenthos_RDA_scores.csv - RDA sample scores of macrozoobenthos in Po - Adriatic Sea river-to-sea system.
MBO_Report_Po_Phytoplankton_RDA_env_vectors.csv - RDA environmental vectors of phytoplankton in Po - Adriatic Sea river-to-sea system.
MBO_Report_Po_Phytoplankton_RDA_scores.csv - RDA sample scores of macrozoobenthos in Po - Adriatic Sea river-to-sea system.



##### Columns:

1. Environmental vectors
RDA1 – Coordinate of the variable along RDA axis 1.
RDA2 – Coordinate of the variable along RDA axis 2.
Variable – Name of the environmental variable.

2. Sample scores 
SampleID – Unique identifier of the sample (site and year).
RDA1 – Coordinate of the sample along RDA axis 1.
RDA2 – Coordinate of the sample along RDA axis 2.
NameID – Site Name.
Habitat – Habitat classification (Freshwater, Transitional, Coastal, Marine).



##### Source Data:

The raw data used in this study are not redistributed here.

They originate from:

Dataset: Das Fachinformationssystem (FIS) der FGG Elbe
Institution/Creators/Authors: River Basin Community Elbe
Repository/Publisher: www.elbe-datenportal.de
DOI/Link: https://www.elbe-datenportal.de/FisFggElbe/content/start/BesucherUnbekannt.action
Citation: © FGG Elbe; data source www.fgg-elbe.de, accessed 04/2024, 05/2024, 07/2024, 08/2024, 09/2024, 04/2025, 08/2025
Additional Information: time period - year 2000 - 2023; location - Brunsbüttelkoog – Strom-km 694,0, Brunsbüttelkoog – Strom-km 694,1, Cuxhaven (Strom-km 725,2), Dommitzsch, linkes Ufer (Strom-km 172,6), FB Dommitzsch, FB Klietznik / Tangermünde, FB Schmilka, FB Schnackenburg / Gorleben, FB Wittenberg, Grauerort (Strom-km 660,6), Magdeburg, linkes Ufer (Strom-km 318,1), Schmilka, rechtes Ufer (Strom-km 3,9), Schnackenburg (Strom-km 474,5), Seemannshöft (Strom-km 628,9), Tonne 117 (Lühemündung) – Strom-km 645,5, Wittenberg, Strommitte (Strom-km 213,8); type - Makrozoobenthos, Fischfauna, Schadstoffe in Wasserphase, allgemeine Gewässergüte; sample type - Makrozoobenthos: Poolprobe; Fischfauna: als Gesamtanzahl; Schadstoffe in Wasserphase/ allgemeine Gewässergüte: Wasser - Gesamtprobe, Einzelprobe, Intervallmessungen

Dataset: Artenliste Phytoplankton Elbe 2004-2023
Institution/Creators/Authors: Saxon State Office for Environment, Agriculture and Geology (LfULG)
Repository/Publisher: n/a
DOI/Link: n/a
Citation: © Sächsischen Landesamtes für Umwelt, Landwirtschaft und Geologie, Datenerhebung: Betriebsgesellschaft für Umwelt und Landwirtschaft, 09/12/2024

Dataset: Phytoplankton Daten des Datenportal des Gewässerkundlichen Landesdienstes des Landesbetrieb für Hochwasserschutz und Wasserwirtschaft Sachsen-Anhalt (LHW)
Institution/Creators/Authors: State Agency for Flood Protection and Water Management of Saxony-Anhalt (LHW)
Repository/Publisher: Gewässerkundlicher Landesdienst - Datenportal
DOI/Link: https://gld.lhw-sachsen-anhalt.de/
Citation: © Landesbetrieb für Hochwasserschutz und Wasserwirtschaft Sachsen-Anhalt (LHW), 09/10/2024
Additional Information: time period - year 2000 - 2022; location - Magdeburg & Wittenberg; type - Fließgewässer Phytoplankton

Dataset: Daten des Niedersächsischen Landesbetrieb für Wasserwirtschaft, Küsten- und Naturschutz (NLWKN)
Institution/Creators/Authors: Lower Saxony Water Management, Coastal and Nature Protection Agency (NLWKN)
Repository/Publisher: n/a
DOI/Link: n/a
Citation: © 2024 Niedersächsischer Landesbetrieb für Wasserwirtschaft, Küsten- und Naturschutz (NLWKN), www.nlwkn.niedersachsen.de/opendata
License: Data licence Germany – attribution – version 2.0 (DL-DE->BY-2.0), www.govdata.de/dl-de/by-2-0

Dataset: Data collection on phytoplankton from the Elbe River near Hamburg [dataset]
Institution/Creators/Authors: Institute for Hygiene and Environment (HU), Diana Rieger, Regine Redelstein 
Repository/Publisher: PANGAEA
DOI/Link: https://doi.pangaea.de/10.1594/PANGAEA.980525
Citation: Rieger, Diana; Redelstein, Regine (2025): Data collection on phytoplankton from the Elbe River near Hamburg [dataset]. PANGAEA; https://doi.pangaea.de/10.1594/PANGAEA.980525
License: Creative Commons Attribution 4.0 International (CC-BY-4.0)

Dataset: Daten des Landesamt für Umwelt des Landes Schleswig-Holstein
Institution/Creators/Authors: State Office for the Environment of Schleswig-Holstein (LfU)
Repository/Publisher: n/a
DOI/Link: n/a
Citation: © Landesamt für Umwelt des Landes Schleswig-Holstein, 28/06/2024 und 26/07/2024
License: Data licence Germany – attribution – version 2.0 (DL-DE->BY-2.0), www.govdata.de/dl-de/by-2-0

Dataset: Dataset on bottom trawl surveys
Institution/Creators/Authors: International Council for the Exploration of the Sea (ICES)
Repository/Publisher: dataXplorer of International Council for the Exploration of the Sea (ICES)
DOI/Link: https://data.ices.dk, https://datras.ices.dk/Data_products/Download/Download_Data_public.aspx
Citation: ICES data portal, Dataset on bottom trawl surveys, 12/06/2025 of data from https://data.ices.dk. ICES, Copenhagen
License: Creative Commons (CC BY 4.0, Attribution 4.0 International)
Additional Information: time period - year 2000 - 2022; area - NS-IBTS (North Sea International Bottom Trawl Survey); type - Unaggregated trawl and biological information

Dataset: Crustaceans and fish abundances and species at and around artificially introduced tetrapod fields in the southern North Sea 2009
Institution/Creators/Authors: Alfred Wegener Institute, Helmholtz Centre for Polar and Marine Research (AWI), Stephanie Wehkamp, Philipp Fischer  
Repository/Publisher: PANGAEA
DOI/Link: https://doi.org/10.1594/PANGAEA.821909
Citation: Wehkamp, Stephanie; Fischer, Philipp (2013): Crustaceans and fish abundances and species at and around artificially introduced tetrapod fields in the southern North Sea 2009 [dataset]. PANGAEA, https://doi.org/10.1594/PANGAEA.821909
License: Creative Commons Attribution 3.0 Unported (CC-BY-3.0)

Dataset: Crustaceans and fish abundances and species at and around artificially introduced tetrapod fields in the southern North Sea, 2010-2011
Institution/Creators/Authors: Alfred Wegener Institute, Helmholtz Centre for Polar and Marine Research (AWI), Stephanie Wehkamp, Philipp Fischer
Repository/Publisher: PANGAEA
DOI/Link: https://doi.org/10.1594/PANGAEA.821915
Citation: Wehkamp, Stephanie; Fischer, Philipp (2013): Crustaceans and fish abundances and species at and around artificially introduced tetrapod fields in the southern North Sea, 2010-2011 [dataset]. PANGAEA, https://doi.org/10.1594/PANGAEA.821915
License: Creative Commons Attribution 3.0 Unported (CC-BY-3.0)

Dataset: Rete Regionale per la qualità ambientale acque superficiali fluviali. Dati 2010-2024 
Institution/Creators/Authors: Regional Agency for Prevention, Environment and Energy of Emilia-Romagna, Silvia Franceschini
Repository/Publisher: Portale Open Data di Arpae
DOI/Link: https://dati.arpae.it/dataset/rete-regionale-per-la-qualita-ambientale-acque-superficiali-fluviali-dati-2010-2025
Additional Information: time period - year 2010 - 2024

Dataset: Benthic diatoms, Macroinvertebrates
Institution/Creators/Authors: Regional Agency for Prevention, Environment and Energy of Emilia-Romagna, Veronica Menna, Alessandra Agostini
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Macroinvertebrates, Phytoplankton, Environmental data
Institution/Creators/Authors: Regional Agency for Prevention, Environment and Energy of Emilia-Romagna - Struttura Oceanografica Daphne, Silvia Pigozzi, Cristina Mazziotti
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Macroinvertebrates
Institution/Creators/Authors: University of Ferrara, Department of Chemical, Pharmaceutical and Agricultural Sciences - DOCPAS, Michele Mistri 
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Environmental data
Institution/Creators/Authors: National Research Council of Italy - Institute of Marine Sciences (CNR-ISMAR), Francesco Riminucci, Caterina Bergami
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Macrozoobenthic Fauna from the Danube–Black Sea System (freshwater, transitional and marine waters) (2003 - 2023)
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Adrian Teaca, Tatiana Begun, Ana-Bianca Pavel 
Repository/Publisher: n/a
DOI/Link: n/a
Citation: Teaca Adrian, Begun Tatiana, Pavel Ana-Bianca. Dataset: Macrozoobenthic Fauna from the Danube–Black Sea System (freshwater, transitional and marine waters) (2003 - 2023)

Dataset: Hydrochemistry in the Romanian shelf waters - July 2023 (PN 23 30 01 03 - National Project) [Dataset]
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Lucian Vasiliu, Naliana Lupascu, Andra Stefania Bucse, Daniela Florea, Florina Radulescu, Sorin Vasile Balan
Repository/Publisher: Zenodo
DOI/Link: https://doi.org/10.5281/zenodo.16744635
Citation: Vasiliu, D. L., Naliana, L., Bucse, A. S., Daniela, F., Radulescu, F.& Balan, S. V. (2025). Hydrochemistry in the Romanian shelf waters - July 2023 (PN 23 30 01 03 - National Project) [Dataset]. Zenodo. https://doi.org/10.5281/zenodo.16744635

Dataset: Hydrochemistry in the Romanian shelf waters - July 2024 (PN 23 30 01 03 - National Project) [Dataset]
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Lucian Vasiliu, Florina Radulescu, Andra Stefania Bucse, Sorin Vasile Balan, Naliana Lupascu, Florina Macau, Daniela Florea
Repository/Publisher: Zenodo
DOI/Link: https://doi.org/10.5281/zenodo.17376980
Citation: Vasiliu, D. L., Radulescu, F., Bucse, A. S., Balan, S. V., Naliana, L., Macau, F.& Daniela, F. (2025). Hydrochemistry in the Romanian shelf waters - July 2024 (PN 23 30 01 03 - National Project) [Dataset]. Zenodo. https://doi.org/10.5281/zenodo.17376980

Dataset: Water column and surface sediment chemistry in the Romanian inner shelf, 2022 (H2020 BRIDGE-BS project) [Dataset]
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar & Institutul Naţional de Cercetare Dezvoltare Marină Grigore Antipa, Dan Lucian Vasiliu, Valentina Coatu, Adrian Teaca, Andra Oros, Andra Bucse, Oana Vlas, Nicoleta-Alexandra Damir, Florina Radulescu
Repository/Publisher: Zenodo
DOI/Link: https://doi.org/10.5281/zenodo.15114215
Citation: GeoEcoMar & Institutul Naţional de Cercetare Dezvoltare Marină Grigore Antipa. (2025). Water column and surface sediment chemistry in the Romanian inner shelf, 2022 (H2020 BRIDGE-BS project) [Dataset]. Zenodo. https://doi.org/10.5281/zenodo.15114215

Dataset: Water physico-chemical parameters (MN101_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2012
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Vasiliu, Dan Secrieru, Sorin Balan
Repository/Publisher: GeoEcoMar archive
DOI/Link: n/a
Citation: Vasiliu Dan, Secrieru Dan, Balan Sorin. Dataset: Water physico-chemical parameters (MN101_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2012

Dataset: Water physico-chemical parameters (MN107_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2013
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Vasiliu, Dan Secrieru, Sorin Balan
Repository/Publisher: GeoEcoMar archive
DOI/Link: n/a
Citation: Vasiliu Dan, Secrieru Dan, Balan Sorin. Dataset: Water physico-chemical parameters (MN107_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2013

Dataset: Water physico-chemical parameters (MN114_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2013
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Vasiliu, Dan Secrieru, Sorin Balan
Repository/Publisher: GeoEcoMar archive
DOI/Link: n/a
Citation: Vasiliu Dan, Secrieru Dan, Balan Sorin. Dataset: Water physico-chemical parameters (MN114_PN 09 41 01 01/HYPOX - Geoecological monitoring of the Romanian shelf) (GeoEcoMar archive), 2013

Dataset: Geochemistry (MN151_PN16450101), 2016
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dan Vasiliu, Dan Secrieru, Sorin Balan 
Repository/Publisher: n/a
DOI/Link: n/a
Citation: Vasiliu Dan, Secrieru Dan, Balan Sorin. Dataset: Geochemistry (MN151_PN16450101), 2016

Dataset: Chemical Parameters of Danube Water Collected During the April–May 2010 Campaign (PN 09 41 03 05)
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dumitru Grosu
Repository/Publisher: n/a
DOI/Link: n/a
Citation: Grosu Dumitru. Dataset: Chemical Parameters of Danube Water Collected During the April–May 2010 Campaign (PN 09 41 03 05)

Dataset: Chemical Elements in Danube Water Samples, June 2003 (GeoEcoMar archive)
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Dumitru Grosu
Repository/Publisher: GeoEcoMar archive
DOI/Link: n/a
Citation: Grosu Dumitru. Dataset: Chemical Elements in Danube Water Samples, June 2003 (GeoEcoMar archive)

Dataset: Phytoplankton Danube 2009 - 2012 (GeoEcoMar archive) Geoecological monitoring program
Institution/Creators/Authors: National Institute for Research and Development on Marine Geology and Geo-ecology – GeoEcoMar, Marian-Traian Gomoiu, Ion Carausu 
Repository/Publisher: GeoEcoMar archive
DOI/Link: n/a
Citation: Gomoiu M-T., Carausu I. Dataset: Phytoplankton Danube 2009 - 2012 (GeoEcoMar archive) Geoecological monitoring program 

Dataset: Fish, Macrophytes, Environmental data
Institution/Creators/Authors: ICPDR – International Commission for the Protection of the Danube River
Repository/Publisher: ICPDR – International Commission for the Protection of the Danube River, Joint Danube Survey
DOI/Link: https://data.danubesurvey.org/, https://www.icpdr.org/library/publications/tnmn-yearbooks                                                       
Citation: © ICPDR - International Commission for the Protection of the Danube River
Additional Information: time period - year 2000 - 2021; location - Romanian Danube

Dataset: Fish, Macrophytes, Benthic diatoms, Phytoplankton, Macrozoobenthos (RO142900, RO14270)
Institution/Creators/Authors: National Administration "Romanian Waters" (ANAR), Nicoleta Rotaru
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Benthic macroinvertebrates and environmental data collected by Port of Sevilla
Institution/Creators/Authors: Port of Sevilla, Antonio Bejarano
Repository/Publisher: n/a
DOI/Link: n/a

Dataset: Fish and environmental data by Universidad de Sevilla
Institution/Creators/Authors: Universidad de Sevilla, Juan Miguel Miró, César Megina, Íñigo Donázar, Jose Carlos García
Repository/Publisher: n/a
DOI/Link: n/a

Please refer to the original source for access conditions and licensing.



##### Data Processing:

The results in this repository were generated using the following steps:

Data acquisition from the original source.
Preprocessing (cleaning, filtering, normalization).
Analysis.
Aggregation and summarization of outputs.

For full details, see the https://codebase.helmholtz.cloud/marco-bolo-task-3.3/deliverable-3.3 directory and accompanying documentation.



##### To reproduce these results:

Obtain the original dataset from the source listed above.
Run the scripts in the https://codebase.helmholtz.cloud/marco-bolo-task-3.3/deliverable-3.3 directory.



##### Limitations:

The shared results are derived and aggregated, and cannot be used to reconstruct the original dataset.
Any interpretations should consider the limitations of the original data source.



##### License:

This data is licensed under the Creative Commons Attribution 4.0 International license.

Note: This license applies only to the materials provided here.



##### Citation:

If you use these results, please cite:


Hufnagel, L., Kamjunke, N., Weitere, M., Bergami, C., Adamescu, C. M., Arhire, G. L., Mureşan, M., Madon, B., Keuter, S., Peredo Arce, A., Sanders, T., Haase, P. (2026). Dataset of Deliverable 3.3 of EU Horizon Europe project MARCO-BOLO: Report on chains of effects and connectivity points and how these effects are reflected by biodiversity assessment results along the Land-River-Sea continuum. Zenodo. https://doi.org/10.5281/zenodo.21455121


And also cite the original datasets.



##### Contact:

Lili Hufnagel: lili.hufnagel@ufz.de
Norbert Kamjunke: nobert.kamjunke@ufz.de
Markus Weitere: markus.weitere@ufz.de